How Homology Is Different from Convergent Evolution Biology
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How Homology Is Different from Convergent Evolution BiologyQuestion 1Pine trees that are too tall or too short do not do as well as pine trees that are averagein height. The short trees do not get as much light as tall or average trees. The tall treesare more likely to break off in storms. Tell how this is an example of stabilizing selection.Be sure to define stabilizing selection in your answer.Your response must be at least 75 words in length.Question 2There is a moth in England called the peppered moth. Before Britain's industrialrevolution, these moths were usually salt and pepper colored. Because of their coloring,they blended in well with the tree trunks on which they tended to rest. The coloringhelped them hide from the birds that ate them. During the British industrial revolution,industry expelled a lot of soot from the burning of coal into the environment. This sootdarkened the tree trunks, and it was noted that black-colored moths were becomingpredominant. The idea is that with soot in the environment, black-colored moths faredbetter than light-colored moths. There is some debate as to whether this is actually thecase or not, but for the sake of this question, let's assume it is.the observations were in the text book i sent you under 12.2. I copied the text here butit’s not really reader friendly in this format. Can you please adjust question #2appropriately? Thanks in advance12.2 Natural Selection Causes Evolution In the Origin of Species, Charles Darwin putforth two major ideas: the theory of common descent (Chapter 11) and the theory ofnatural selection. Darwin’s presentation of the theory of common descent—that allspecies living today appear to have descended from a single ancestor—was thoroughand convincing.Within 20 years of the publication of his book, the theory of common descent had beenaccepted by most scientists. However, it was another 60 years before the scientificcommunity accepted Darwin’s theory of natural selection, which explains in large parthow organisms evolved from a common ancestor to become the great variety we seetoday.Darwin proposed that through the process of natural selection, the physical orbehavioral traits of organisms that lead to increased survival or reproduction becomecommon within their population, while less favorable traits are lost. The changesaccumulating within populations via natural selection can lead to the development ofnew species.Darwin reasoned that the process of natural selection is an inevitable consequence ofthe competition for survival among variable individuals in a population. Today, naturalselection is considered one of the most important causes of evolution (although others,such as the processes of genetic drift and sexual selection as described in Chapter 13,also cause populations to change over time).Darwin’s Observations The theory of natural selection is elegantly simple. It is aninference based on four general observations: 1. Individuals within Populations VaryObservations of groups of humans support this statement—people do come in anenormous variety of shapes, sizes, colors, and facial features. It may be less obviousthat there is variation in nonhuman populations as well.For example, in a litter of gray wolves born to a single female, individuals may vary incoat color, while in a field of flowers, one plant may bloom earlier than others (Figure12.6). We can add all kinds of less obvious differences to this visible variation; forexample, the amount of caffeine produced in the seeds of a coffee plant varies amongindividuals in a wild population. Each different type of individual in a population is calleda variant. Visualize This Under what conditions might it be an advantage for anindividual plant to bloom earlier than other nearby flowers? Figure 12.6 Observation 1:Individuals within populations vary. (a) Gray wolves vary in coat color, even within asingle litter of animals. (b) Flowers may vary in blooming time, with some individualplants blooming much earlier than others of the same species. Figure 12.6 FullAlternative Text 2. Some of the Variation among Individuals Can Be Passed on to TheirOffspring Although Darwin did not understand how it occurred, he observed manyexamples of the general resemblance between parents and offspring. He also noticedthat people took advantage of the inheritance of variation in other species.Pigeon fanciers in Darwin’s time clearly recognized the inheritance of variation; theycould see, for instance, that pigeons with neck ruffs were more likely to produceoffspring with neck ruffs than were pigeons without ruffs. Thus, when enthusiastswanted to produce a ruffed variety of pigeon, they encouraged breeding among thebirds with this trait (Figure 12.7). Darwin hypothesized that offspring tend to have thesame characteristics as their parents in natural populations as well. Figure 12.7Observation 2: Some of the variation among individuals can be passed on to theiroffspring.Darwin noted that breeders could create flocks of pigeons with fantastic traits by usingas parents of the next generation only those individuals that displayed these traits. Forseveral decades after the Origin of Species was published, the observation that somevariations were inherited was the most controversial part of the theory of naturalselection. Because scientists could not adequately explain the origin and inheritance ofvariation, many were unwilling to accept that natural selection could be a mechanism forevolutionary change. When Gregor Mendel’s work on inheritance in pea plants (Chapter8) was rediscovered in the 1900s, the mechanism for this observation becameclear—natural selection operates on genetic variation that is passed from onegeneration to the next. 3.Populations of Organisms Produce More Offspring than Will Survive This observation isclear to most of us—the trees in the local park make literally millions of seeds everysummer, but only a small fraction of these survive to germinate, and only a few of theseedlings live for more than a year or two. In the Origin of Species, Darwin gave agraphic illustration of the difference between offspring production and survival. In hisexample, he used elephants, animals that live long lives and are very slow breeders.A female elephant does not begin breeding until age 30, and she produces about 1 calfevery 10 years until around age 90. Darwin calculated that even at this very low rate ofreproduction, if all the descendants of a single pair of African elephants survived andlived full, fertile lives, after about 500 years their family would have more than 15 millionmembers (Figure 12.8)—many more than can be supported by all the available foodresources on the African continent! VisualizeThis Predict what will happen to the elephant population as a result of limited resourceavailability when Generation 4 is produced. Figure 12.8 Observation 3: Populations oforganisms produce more offspring than will survive. Even slow-breeding animals likeelephants are capable of producing huge populations relatively quickly. Figure 12.8 FullAlternative Text 4. Survival and Reproduction Are Not Random In other words, thesubset of individuals that survives long enough to reproduce is not an arbitrary group.Some variants in a population have a higher likelihood of survival and reproduction thanother variants do; that is, there is differential survival and reproduction amongindividuals in the population.The survival and reproduction of one variant compared with others in the samepopulation is referred to as its relative fitness. Traits that increase an individual’s relativefitness in a particular environment are called adaptations. Individuals with adaptations toa particular environment are more likely to survive and reproduce than are individualslacking such adaptations; in other words, these individuals have higher relative fitness.Darwin referred to the results of differential survival and reproduction as naturalselection.Adaptations are “naturally selected” in the sense that individuals possessing themsurvive and contribute offspring to the next generation. Although Darwin used the wordselection, which implies some active choice, natural selection is a passive process thatis simply determined by differences among individuals and their success in theirparticular environment. For example, among the birds called medium ground finchesliving on an island in the Galápagos archipelago, scientists have observed that whenrainfall is scarce, a large bill is an adaptation that can be observed.The large bill can be explained because birds with this attribute are able to crack openlarge, tough seeds—the only food available during severe droughts. As shown in Figure12.9, the 90 survivors of a 1977 drought had an average bill depth that was 6% greaterthan the average bill depth of the original population of 751 birds. In theseenvironmental conditions, a large bill increases survival. Working with Data Use thegraph to determine how the total population size of ground finches changed between1976 and 1978. Figure 12.9 Observation 4: Survival and reproduction are not random.The pale purple curve summarizes bill depth in ground finches on Daphne Island in theGalápagos in 1976. The dark purple curve below it represents the population in 1978,after the drought of 1977. These data indicate that survivors of the drought had a largeraverage bill depth than the predrought population. The change in the population’saverage bill size occurred because finches with larger-than-average bills had higherfitness than did small-billed birds during the drought. Figure 12.9 Full Alternative TextAdaptations are not only traits that increase survival.Any trait possessed by an individual that increases the number of offspring it producesrelative to other individuals in a population is also an adaptation. For example, flowers ina meadow may have a relatively limited number of potential insect pollinators. Morepollinator visits generally result in more seeds being produced by a single flower, so anytrait that increases a flower’s attractiveness to pollinators, such as a brighter color orgreater nectar production, should be favored by natural selection (Figure 12.10). Figure12.10 Adaptations are not about survival only. Variations that increase a flower’sattractiveness to a pollinator can increase its reproductive success by increasing thenumber of seeds it produces. Darwin’s Inference: Natural Selection Causes EvolutionBased on his observations, Darwin reasoned that the result of natural selection is thatinherited variations that are favorable within a given environment tend to increase infrequency in a population over time, while variations that are unfavorable within a givenenvironment tend to be lost within the population. In other words, adaptations becomemore common in a population as the individuals who possess them contribute largernumbers of their offspring to the succeeding generation. Natural selection results in achange in the traits of individuals in a population over the course of generations—thatis, evolution. Although there are other factors, such as genetic drift and migration ofindividuals, that can cause populations to evolve over time, natural selection is the onlyforce that can lead to the adaption of a population to its environment. It is a testament tothe power of the theory of natural selection that today it seems self-evident to us. Butthe theory of natural selection only became so powerful after it was tested and shown towork—in nature—in the manner Darwin described. Natural selection proved such apowerful idea that it has influenced how we think about many phenomena, from thesuccess of particular brands of soft drinks to the relationships among nations. Naturalselection also explains the emergence of XDR-TB. Testing Natural Selection Darwinproposed a scientific explanation of how evolution occurs, and like all good hypotheses,it needed to be tested. All of the tests described next illustrate that natural selection isan effective mechanism for evolutionary change. Artificial Selection Selection imposedby human choice is called artificial selection. It is artificial in the sense that humansdeliberately control the survival and reproduction of individual plants and animals tochange the characteristics of the population. Individuals with preferred traits arepermitted to breed, whereas those that lack preferred traits are not allowed to breed.The fancy pigeons that Darwin studied arose by artificial selection, and the great varietyof domestic dogs we see today also resulted from this process. In each case, differentbreeds evolved through selection by breeders for various traits (Figure 12.11). Theseexamples demonstrate that differential survival and reproduction change thecharacteristics of populations. However, because of the direct intervention of humanson the survival and reproduction of these organisms, artificial selection is not exactlyequivalent to natural selection. Can change in populations occur without direct humanintervention? Visualize This What would this sequence of changes look like if theselection was for dogs with a particular behavioral trait, for instance, pointing at a preyanimal? Figure 12.11 Artificial selection can cause evolution. When breeders selectdogs with certain traits to produce the next generation of animals, they increase thefrequency of that trait in the population. Over generations, the trait can become quiteexaggerated. Dachshunds are descendants of dogs that were selected for theproduction of very short legs. Figure 12.11 Full Alternative Text Natural Selection inthe Lab Another test of the effectiveness of natural selection is to examine whetherpopulations living in artificially manipulated laboratory environments change over time.An example of this kind of experiment is one performed on fruit flies placed inenvironments containing different concentrations of alcohol. High concentrations ofalcohol cause cell death in fruit flies. Many organisms, including fruit flies and humans,produce enzymes that metabolize alcohol—that is, they break it down, extract energyfrom it, and modify it into less-toxic chemicals. There is variation among fruit flies in therate at which they metabolize alcohol. In a typical laboratory environment, most fliesprocess alcohol relatively slowly, but about 10% of the population possesses anenzyme variant that allows those flies to metabolize alcohol twice as rapidly as the morecommon variant. In an experiment (Figure 12.12), scientists divided a population of fruitflies into two randomized groups. Initially, these two groups had the same percentage offast and slow alcohol metabolizers. One group of flies was placed in an environmentcontaining typical food sources; the other group was placed in an environmentcontaining the same food spiked with alcohol. After 57 generations, or about two yearsin the laboratory, the percentage of fast alcohol-metabolizing flies in the environmentwith only typical food sources was the same as at the beginning of theexperiment—10%. But after the same number of generations, the percentage of fastalcohol-metabolizing flies in the alcohol-spiked environment was 100%. Because all ofthe flies in this environment were now of the fast alcohol-metabolizing variety, theaverage rate of alcohol metabolism in the population in this environment was muchhigher in generation 57 than in generation 1. The population had evolved. Working withData This data can be represented as a line graph as well as a bar graph. Sketch theline graph. Figure 12.12 Natural selection in laboratory conditions. When fruit flies areplaced in a high-alcohol environment, the percentage of flies that can rapidly metabolizealcohol increases over many generations because of natural selection. In the normallaboratory environment, there is no selection for faster alcohol processing, so theaverage rate of alcohol metabolism does not change. Figure 12.12 Full Alternative TextThe evolution of the fruit flies in this experiment was a result of natural selection. In anenvironment where alcohol concentrations were high, individuals that were able tometabolize alcohol relatively rapidly had higher fitness. Because they lived longer andwere less affected by alcohol, the fast alcohol-metabolizing flies left more offspring thanthe slow alcohol- metabolizing flies did. Thus, each generation had a higher frequencyof fast alcohol-metabolizing individuals than the previous generation did. After manygenerations, flies that could rapidly metabolize alcohol predominated in the population.Selection can change populations in highly regulated laboratory environments. But doesit have an effect in natural wild populations? Natural Selection in Wild Populations Theevolution of M. tuberculosis from being susceptible to antibiotics to being resistant isone example of natural selection in a wild population; clearly, a change in theenvironment (that is, the introduction of antibiotics) caused a change in the bacteriapopulation. Dozens of other pathogens, organisms that cause disease, have becomeresistant to drugs and pesticides in the past 50 years as well. But even these changesmay seem less convincing to some readers because the adaptation is to a human-imposed environmental change. Although studying adaptation to natural environmentalchanges in the field is a significant challenge, the effects of natural selection have alsobeen observed in dozens of wild populations. A classic example of natural selection in anatural setting is the evolution of bill size in Galápagos finches in response to drought(review Figure 12.9). The survivors of the drought tended to be those with the largestbills, which could more easily handle the tough seeds that were available in the dryenvironment. The survival of this nonrandom subset of birds resulted in a dramaticchange in the next generation. The population of birds that hatched from eggs in1978—the descendants of the drought survivors—had an average bill depth 4 to 5%larger than that of the predrought population. A more recent example of naturalselection causing evolution has occurred in the past few decades on the eastern coastof the United States, where an invasive Asian crab species is wreaking havoc on nativemussels. But one species, the blue mussel, has quickly evolved the ability to thicken itsshell when it grows in the presence of the Asian crab, thwarting their attacks (Figure12.13). Scientists at the University of New Hampshire were able to demonstrate that thiswas an evolutionary change by comparing blue mussel populations in regions invadedby the Asian crab with those in more northerly waters, where the Asian crab cannotsurvive. These researchers demonstrated that while both populations of musselsthicken their shells in response to the presence of native crabs, only the mussels thathad been living with the Asian crabs responded to the presence of this species. Clearly,natural selection for individual mussels that could recognize this new species of crab asa predator had caused a change in the mussel population. Figure 12.13 Naturalselection in the wild. (a) Asian shore crabs are recent invaders to the east coast;because they were unfamiliar predators to the native mussels, they initially devastatedmussel beds. (b) Some populations of blue mussels have evolved the ability torecognize the presence of the crabs and respond the way they do to native predators,by adding layers to their shells. Got It? Two of Darwin’s observations that led to thetheory of natural selection are that organisms in a population from each other and thattraits can be on their offspring. The fitness of an individual is defined as its success inand compared with other individuals in the same population. An adaptation is a trait thatincreases an individual’s relative to others in the population who do not have the trait.When humans manipulate the environment and cause the evolution of traits in apopulation of domestic animals or plants it is called . The evolution of rapid alcoholmetabolism in a population of fruit flies in a lab occurred in the presence of high alcohollevels could occur because individuals in their rate of alcohol metabolism.In your own words, explain the concepts from the four observations discussed in 12.2using the moth as an example. In other words, how does the moth illustrate the firstobservation, the second observation, etc.?Your response must be at least 200 words in length.Question 3Explain how homology is different from convergent evolution and give examples. Brieflydefine homology and convergent evolution in your explanation.How Homology Is Different from Convergent Evolution Biology
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How Homology Is Different from Convergent Evolution Biology